Okay , I'm coming back to option A (front vs rear pairs)
I have found out the term for this is "staggered jetting" , apparently it is commony used on V fours to make the rear cylinders run cooler (richer mix)
It is also used to correct for differing intake lengths or differing cooling on the exhaust headers.
I am not aware of any difference on St1100 air intakes, if we consider the cooling and exhaust headers then the difference if any would be front to rear
From my perspective it is possible to read the parts fiche diagram such that the front pair of needles are the same (and the rear pair are the same but different from the front pair)